Down below you will find the full bibliography of my recent studies

Technical Papers


Hendrickx C, Bell PR, Ezcurra M, Cuesta E, Milner AR, Xu X, O’Connor JK, Loewen M, Sereno PC, Mateus O. 2022. Morphology and distribution of epidermal structures (excluding feathers) in non-avian theropod dinosaurs. Biological Reviews (Cambridge).


Atterholt JA, Poust AW, Erickson GM, O’Connor JK. 2021. Intraskeletal osteohistovariability reveals complex growth strategies in a Late Cretaceous enantiornithine. Frontiers in Earth Science, 9:640220.

Clark AD, O’Connor JK. 2021. Exploring the ecomorphology of two Cretaceous enantiornithines with unique pedal morphology. Frontiers in Ecology and Evolution, 15:654156.

Kiat Y, Pyle P, Balaban A, O’Connor JK. 2021. Reinterpretation of purported molting evidence in the Thermopolis Archaeopteryx. Communications Biology, 4:837

Liu S-M, Li Z-H, Bailleul AM, Wang M, O’Connor JK. 2021. Investigating possible gastroliths in a referred specimen of Bohaiornis guoi (Aves: Enantiornithes). Frontiers in Earth Science, 9:635727.

O’Connor JK, Stidham TA, Harris JD, Lamanna M, Bailleul AM, Hu H, Wang M, You H-L. 2021. Avian skulls represent a diverse ornithuromorph fauna from the Lower Cretaceous Xiagou Formation, Gansu Province, China. Journal of Systematics and Evolution.

Stidham TA, O’Connor JK. 2021. The evolutionary and functional implications of the unusual quadrate of Longipteryx chaoyangensis (Avialae: Enantiornithes) from the Cretaceous Jehol Biota of China. Journal of Anatomy, 239(5), 1066-1074.

Talori YS, Zhao J-S, O’Connor JK. 2021. Kinematics of wings from Caudipteryx to modern birds.Proceedings of the Institution of Mechanical Engineers, Part C: Journal of Mechanical Engineering Science.

Wang M, O’Connor JK, Zhao T, Pan Y-H, Zheng X-T, Wang X-L, Zhou Z-H. 2021. An Early Cretaceous enantiornithine bird with a pintail. Current Biology, 31(21), 4845-4852.

Wu Q, O’Connor JK, Li Z-H, Bailleul AM. 2021. Cartilage on the furculae of living birds and the extinct bird Confuciusornis: a preliminary analysis and implications for flight style inferences in Mesozoic birds. Vertebrata Palasiatica, 59(2): 110-124.

Wu Q, Bailleul AM, Li Z-H, O’Connor JK, Zhou Z-H. 2021. Osteohistology of the scapulocoracoid of Confuciusornis and preliminary analysis of the shoulder joint in Aves. Frontiers in Earth Science, 9: 617124.

Xu L, Buffetaut E, O’Connor JK, Zhang X-L, Jia S-H, Zhang J-M, Chang H-L, Tong H-Y. 2021. A new, remarkably preserved, enantiornithine bird from the Late Cretaceous of Henan (central China) and convergent evolution between enantiornithines and modern birds. Geological Magazine, 158(11), 2087-2094.


Hu H, O’Connor JK, Wang M, McDonald PG, Wroe S, Zhou Z-H. 2020. New anatomical information of bohaiornithid Longusunguis confirms plesiomorphic diapsid skull retained in Enantiornithes. Journal of Systematic Palaeontology, In press.

O’Connor JK, Falk AR, Wang M, Zheng X-T. 2020. First report of immature feathers in juvenile enantiornithines from the Early Cretaceous Jehol avifauna. Vertebrata Palasiatica, 58(1). 24-44.

Xing L-D, O’Connor JK, Schmitz L, Chiappe LM, McKellar RC, Yi Q-R, Li G. 2020. Hummingbird-sized dinosaur from the cretaceous period of myanmar. Nature 579, 245-249. (corresponding author)

Xing L-D, McKellar RC, O’Connor J. 2020. An unusually large bird wing in mid-cretaceous burmese amber. Cretaceous Research, 110: 104412.


O’Connor JK, Zhou Z-H. 2019. The evolution of the modern avian digestive system – insights from paravian fossils from the Yanliao and Jehol biotas. Palaeontology.

O’Connor JK. 2019. The trophic habits of early birds. Palaeogeography, Palaeoclimatology, Palaeoecology 513, 178-195

O’Connor JK, Zheng X-T, Dong L-P, Wang X-L, Wang Y, Zhang X-M, Zhou Z-H. 2019. Microraptor with ingested lizard suggests non-specialized digestive function. Current Biology, 29: 1-7.

Bailleul AM, Li Z-H, O’Connor JK, Zhou Z-H. 2019. Origin of the avian predentary and evidence of a unique form of cranial kinesis in Cretaceous ornithuromorphs. Proceedings of the National Academy of Sciences U.S.A., in press.

Bailleul AM, O’Connor JK, Schweitzer MH. 2019. Dinosaur paleohistology: review, trends and new avenues of investigation. PeerJ 7: e7764. 

Bailleul AM, O’Connor JK, Zhang S-K, Li Z-H, Wang Q, Lamanna M, Zhu X-F, Zhou Z-H. 2019. An Early Cretaceous enantiornithine (Aves) preserving an unlaid egg and probable medullary bone. Nature Communications, 10(1275): 1-10.

Falk AR, O’Connor JK, Wang M, Zhou Z-H. 2019. On the preservation of the beak in Confuciusornis (Aves: Pygostylia). Diversity, 11(212): 1-8.

Hu H, Sansalone G, Wroe S, McDonald PG, O’Connor JK, Li Z-H, Xu X, Zhou Z-H. 2019. Evolution of the vomer and its implications for cranial kinesis in Paraves. Proceedings of the National Academy of Sciences U.S.A., 201907754

Pan YH, Zheng W-X, Sawyer RH, Penninngton M, Zheng X-T, Wang X-L, Wang M, Hu L, O’Connor JK, Zhao T, Li Z-H, Schroeter E, Wu F-X, Xu X, Zhou Z-H, Schweitzer MH. 2019. The molecular evolution of feathers with direct evidence from fossils. Proceedings of the National Academy of Sciences U.S.A., 116(8): 3018-3023.

Talori YS, Zhao J-S, Liu Y-F, Lu W-X, Li Z-H, O’Connor JK. 2019. Identification of avian flapping motion from non-volant winged dinosaurs based on modal effective mass analysis. PLoS Computational Biology, 15(5): e1006846.

Wang M, O’Connor JK, Zhou S, Zhou Z-H. 2019. New toothed Early Cretaceous ornithuromorph bird reveals intraclade diversity in pattern of tooth loss. Journal of Systematic Palaeontology.

Wang M, O’Connor JK, Zhou Z-H. 2019. A taxonomical revision of the Confuciusornithiformes (Aves: Pygostylia). Vertebrata Palasiatica, 57(1), 1-37

Wang M, O’Connor JK, Xu X, Zhou Z-H. 2019. A new Jurassic scansoriopterygid and the loss of membranous wings in theropod dinosaurs. Nature, 569(7755): 256-259.

Xing L-D, McKellar RC, O’Connor JK, Bai M, Tseng K-W, Chiappe LM. 2019. A fully feathered enantiornithine foot and wing fragment preserved in mid-Cretaceous Burmese amber. Scientific Reports, 9(927): 1-9

Xing L-D, O’Connor JK, Chiappe LM, McKellar RC, Carroll N, Hu H, Bai M, Lei F-M. 2019. A new enantiornithine with unusual pedal proportions found in amber. Current Biology, 29(14): 2396-2401 (co-first author)

Xing L-D, McKellar RC, O’Connor JK, Niu K-C, Mai H-J. 2019. A mid-cretaceous enantiornithine foot and tail feather preserved in Burmese amber. Scientific Reports, 9:15513.


O’Connor JK, Erickson GM, Norell MA, Bailleul AM, Hu H, Zhou Z-H. 2018. Medullary bone in an Early Cretaceous enantiornithine (Aves) and discussion regarding its identification in fossils. Nature Communications 9, 5169

O’Connor JK, Wang X-L, Sullivan C, Wang Y, Zheng X-T, Hu H, Zhang X-M, Zhou Z-H. 2018. First report of gastroliths in the Early Cretaceous basal bird JeholornisCretaceous Research 84, 200-208

Zheng X-T, O’Connor JK, Wang X-L, Wang Y, Zhou Z-H. 2018. Reinterpretation of a previously described Jehol bird clarifies early trophic evolution in the Ornithuromorpha. Proceedings of the Royal Society B: Biological Sciences 285, 20172494.

Xing L-D, O’Connor JK, McKellar RC, Chiappe LM, Bai M, Tseng K-W, Zhang J, Yang H-D, Fang J, Li G. 2018. A flattened enantiornithine in mid-Cretaceous Burmese amber: Morphology and preservation. Science Bulletin 63(4), 235-243.

Wang X-L, O’Connor JK, Maina JN, Pan Y-H, Wang M, Wang Y, Zheng X-T, Zhou Z-H. 2018. Archaeorhynchuspreserving significant soft tissue including probable fossilized lungs. Proceedings of the National Academy of Sciences U.S.A. 115(45): 11555-11560.

Rashid DJ, Surya K, Chiappe LM, Carroll N, Garrett KL, Varghese B, Bailleul AM, O’Connor JK, Chapman SC, Horner JR. 2018. Avian tail ontogeny, pygostyle formation, and interpretation of juvenile Mesozoic specimens. Scientific Reports 8, 9014.

Atterholt JA, Hutchison JH, O’Connor JK. 2018. The most complete enantiornithine from North America and a phylogenetic analysis of the Avisauridae. PeerJ 6, e5910.

Smith-Paredes D, Núñez-León D, Soto-Acuña S, O’Connor JK, Botelho JF, Vargas AO. 2018. Dinosaur ossification centres in embryonic birds uncover developmental evolution of the skull. Nature Evolution & Ecology 2, 1966-1973

Talori YS, Liu Y-F, Zhao J-S, Sullivan C, O’Connor JK, Li Z-H. 2018. Winged forelimbs of the small theropod dinosaur Caudipteryx could have generated small aerodynamic forces during rapid terrestrial locomotion. Scientific Reports 8:17854

Xing L-D, Cockx O, McKellar RC, O’Connor JK. 2018. Ornamental feathers in Cretaceous Burmese amber: resolving the enigma of rachis-dominated feather structure. Journal of Palaeogeography 7(13):1-18


O’Connor JK, Zheng X-T, Hu H, Wang X-L, Zhou Z-H. 2017. The morphology of Chiappeavis magnapremaxillo(Pengornithidae: Enantiornithes) and a comparison of aerodynamic function in Early Cretaceous avian tail fans. Vertebrata Palasiatica 55, 1-18.

Zheng X-T, O’Connor JK, Pan Y-H, Wang X-L, Wang Y, Wang M & Zhou Z-H. 2017. Exceptional preservation of soft tissue in a new specimen of Eoconfuciusornis and its biological implications. National Science Review 4(3): 441-452.

Wang M, O’Connor JK, Pan Y-H, Zhou Z-H. 2017. A bizarre Early Cretaceous enantiornithine bird with unique crural feathers and an ornithuromorph plough-shaped pygostyle. Nature Communications, 8(14141): 1-12.

Wang W, O’Connor JK. 2017 Morphological coevolution of the pygostyle and tail feathers in Early Cretaceous birds. Vertebrata Palasiatica 55(4): 289-314.

Hu H, O’Connor JK. 2017. First species of Enantiornithes from Sihedang elucidates skeletal development in Early Cretaceous enantiornithines. Journal of Systematic Palaeontology 15(11): 909-926. 

Xing L-D, O’Connor JK, McKellar RC, Chiappe LM, Tseng K-W, Li G, Bai M. 2017 A mid-Cretaceous enantiornithine (Aves) hatchling preserved in Burmese amber with unusual plumage. Gondwana Research 49: 264-277.

Wang Y, Hu H, O’Connor JK, Wang M, Xu X, Zhou Z-H, Wang X-L, Zheng X-T. 2017 A previously undescribed specimen reveals new information on the dentition of Sapeornis chaoyangensisCretaceous Research 74, 1-10.

Sullivan C, Xu X, O’Connor JK. 2017. Complexities and novelties in the early evolution of avian flight, as seen in the Mesozoic Yanliao and Jehol Biotas of northeast China. Palaeoworld 26(2): 212-229.


O‘Connor JK, Li D-Q, Lamanna M, Wang M, Harris JD, Atterholt J, You H-L. 2016. A new Early Cretaceous enantiornithine (Aves: Ornithothoraces) from northwestern China with elaborate tail ornamentation. Journal of Vertebrate Paleontology, 36(1): e1054035.

O’Connor JK, Wang X-L, Zheng X-T, Hu H, Zhang X-M, Zhou Z-H. 2016. An enantiornithine with a fan-shaped tail, and the evolution of the rectricial complex in early birds. Current Biology, 26(1): 114-119.

O’Connor JK, Wang M, Hu H. 2016. A new ornithuromorph (Aves) with an elongate rostrum from the Jehol Biota and the early evolution of rostralization in birds. Journal of Systematic Palaeontology 14(11): 939-948.

Wang Y, Wang M, O’Connor JK, Zhou Z-H, Wang X-L, Zheng X-T, Zhang X-M. 2016. A new three dimensionally preserved Jehol enantiornithine bird with ovarian follicles. Journal of Vertebrate Paleontology 36(2): e1054496.

 Botelho JB, Smith-Paredes D,  Soto-Acuña S, O’Connor JK, Palma V, Vargas A. 2016. Molecular development of fibular reduction in birds and its evolution from dinosaurs. Evolution DOI: 10.1111/evo.12882.

Xing L, McKellar RC, Wang M, Bai M, O’Connor JK, Benton MJ, Zhang J, Wang Y, Tseng K, Lockley MG, Li G. 2016. Mummified precocial bird wings in mid-Cretaceous Burmese amber. Nature Communications 7, 12089.

Pan Y-H, Zheng W-X, Moyer A, O’Connor JK, Wang M, Zheng X-T, Wang X-L, Schroeter E, Zhou Z-H, Schweitzer MH. 2016. Molecular evidence of keratin and melanosomes in feathers of the Early Cretaceous bird EoconfuciusornisProceedings of the National Academy of Sciences U.S.A. 113(49): E7900-E7907.


O’Connor JK, Zheng X-T, Sullivan C, Chuong C-M, Wang X-L, Li A, Wang Y, Zhang X-M & Zhou Z-H. 2015. Evolution and functional significance of derived sternal ossification patterns in ornithothoracine birds. Journal of Evolutionary Biology 28: 1550-1567.

O’Connor JK, Zheng X-T, Wang XL, Zhang X-M & Zhou Z-H. 2015. The gastral basket in basal birds and their close relatives: size and possible function. Vertebrata Palasiatica 53: 133-152.

O’Connor JK & Zhou Z-H 2015. Early evolution of the biological bird: perspectives from new fossil discoveries in China. Journal of Ornithology: 1-10; 10.1007/s10336-015-1222-5.

O’Connor JK & Chang H-L. 2015. Hindlimb feathers in paravians: primarily ‘wings’ or ornaments? Biology Bulletin 42: 1-6.

O’Connor JK, Wang M, Zhou S & Zhou Z-H. 2015. Osteohistology of the Lower Cretaceous Yixian Formation ornithuromorph (Aves) Iteravis huchzermeyeriPalaeontologica Electronica 18.2: 1-11.

Brusatte SL, O’Connor JK, Jarvis ED. 2015. The origin and diversification of birds. Current Biology 25: R888–R898.

Hu H, O’Connor JK & Zhou Z-H. 2015. A new species of Pengornithidae (Aves: Enantiornithes) from the Lower Cretaceous of China suggests a specialized scansorial habitat previously unknown in early birds. PLoS ONE 10: e0126791.

Xu X, Zheng X-T, Sullivan C, Zhang F-C, O’Connor JK, et al. 2015. A bizarre Jurassic maniraptoran theropod with preserved evidence of membranous wings. Nature 521: 70-73.

Wang M, Zheng X-T, O’Connor JK, Lloyd GT, Wang X-L, et al. 2015. The oldest record of Ornithuromorpha reveals heterogeneous rates of morphological evolution among Early Cretaceous birds. Nature Communications 6: 6987. 

Wang Y-M, O’Connor JK, Li D-Q, You H-L. 2015. New information on the postcranial skeleton of the Early Cretaceous Gansus yumenensis (Aves: Ornithuromorpha). Historical Biology DOI:10.1080/08912963.2015.1006217

Wang M, Li D-Q, O’Connor JK, Zhou Z-H, You H-L. 2015. Second species of enantiornithine bird from the Lower Cretaceous Changma Basin, northwestern China with implications for the taxonomic diversity of the Changma avifauna. Cretaceous Research 55:56-65.


O’Connor JK, Sullivan C. 2014. Reinterpretation of the Early Cretaceous maniraptoran (Dinosauria: Theropoda) Zhongornis haoae as a scansoriopterygid-like non-avian, and the morphological resemblances between scansoriopterygids and basal oviraptorosaurs. Vertebrata Palasiatica 52(1):3-30. 

O’Connor JK, Wang M, Zheng X-T, Wang X-L, Zhou Z-H. 2014. The histology of two female Early Cretaceous birds. Vertebrata Palasiatica 52(1):112-128. 

O’Connor JK, Wang M, Zheng X-T, Zhou Z-H. 2014. Reply to Foth: [preserved cartilage is rare but not absent – troodontid sternal plates are absent not rare. Proceedings of the National Academy of Sciences.

Zheng X-T, O’Connor JK, Wang X-L, Wang M, Zhang X-M, Zhou Z-H. 2014. On the absence of sternal elements in Anchiornis (Paraves: Troodontidae) and Sapeornis (Aves: Pygostylia) and the complex early evolution of the avian sternum. Proceedings of the National Academy of Sciences 111(38):13900-13905(corresponding author)

Wang X-L, O’Connor JK, Zheng X-T, Wang M, Hu H, Zhou Z-H. 2014. New information on the evolution of rachis dominated tail feathers from a new basal enantiornithine. Biological Journal of the Linnean Society 113:805-819. (corresponding author)

Zhou S, O’Connor JK, Wang M. A new species from an ornithuromorph (Aves: Ornithothoraces) dominated locality of the Jehol Biota. 2014. Chinese Science Bulletin 59(36): 5366-5378. (corresponding author)

Wang M, O’Connor JK, Zhou Z-H. 2014. A new robust enantiornithine bird from the Lower Cretaceous of China with scansorial adaptations. Journal of Vertebrate Paleontology 34(3): 657-671.

O’Connor JK, Averianov A, Zelenkov NV. 2014. A confuciusornithiform (Aves: Pygostylia) -like tarsometatarsus from the Early Cretaceous of Siberia and a discussion of the evolution of avian hindlimb musculature. Journal of Vertebrate Paleontology 34(3): 647-56.

Zheng X-T, O’Connor JK, Huchzermeyer F, Wang X-L, Wang Y, Zhang X-M, Zhou Z-H. 2014. New specimens of Yanornis indicate a piscivorous diet and modern alimentary canal. PlosOne 9(4): e95036. doi:10.1371/journal.pone.0095036(corresponding author)

Zhang S-K, Jin X-S, O’Connor JK, Wang M, Xie J-F. 2014. A new egg with avian egg shape from the Upper Cretaceous of Zhejiang Province, China. Historical Biology DOI: 10.1080/08912963.2014.902451(corresponding author)

Chiappe LM, Zhao B, O’Connor JK, Gao C-L, Habib M, Marugan-Lobon J, Meng Q-J, Cheng X-D. 2014. A new specimen of the early cretaceous bird Hongshanornis longicresta: Insights into the aerodynamics and diet of a basal ornithuromorph. PeerJ 2:e234; DOI 10.7717/peerj.234

Hu H, Zhou Z-H, O’Connor JK. 2014. A subadult specimen of Pengornis and character evolution in Enantiornithes. Vertebrata Palasiatica 52(1):77-97.

Wang M, O’Connor JK, Zelenkov NV. 2014. A new diverse enantiornithine family (Bohaiornithidae fam. Nov.) From the lower cretaceous of china with information from two new species. Vertebrata Palasiatica 52(1):31-76. 

Zhang Y-G, O’Connor JK, Liu D, Meng Q-J, Sigurdsen T, Chiappe LM. 2014. New information on the anatomy of the Chinese Early Cretaceous Bohaiornithidae (Aves: Enantiornithes) from a subadult specimen of Zhouornis haniPeerJ: e407; DOI 10.7717/peerj.407.

Zheng X-T, O’Connor JK, Wang X-L, Zhou Z-H, Zhang X-M. 2014. New information on Hongshanornithidae (Aves: Ornithuromorpha) from a new subadult specimen. Vertebrata Palasiatica 52(2):217-232.

O’Connor JK, Chang H-L. Hindlimb feathers in paravians: primarily ‘wings’ or ornaments? 2014. Zoological Journal93(10):1166-1172 [in Russian]


O’Connor JK, Zhou Z-H. 2013. A redescription of Chaoyangia beishanensis (Aves) and a comprehensive phylogeny of Mesozoic birds. Journal of Systematic Palaeontology 11(7):889-906.

O’Connor JK, Zelenkov NV. 2013. The phylogenetic position of Ambiortus: comparison with Early Cretaceous birds from Asia. Paleontological Journal RAS 47(11):1270–1281.

O’Connor JK, Wang X-L, Sullivan C, Zheng X-T, Tubaro P, Zhang X-M, Zhou Z-H. 2013. The unique caudal plumage of Jeholornis and complex tail evolution in early birds. PNAS 43(110): 17404-17408.

O’Connor JK, Zheng X-T, Wang X-L, Wang Y, Zhou Z-H. 2013. Ovarian follicles shed new light on dinosaur reproduction during the transition towards birds. National Science Review 1(1): 15-17.

O’Connor JK, Zhang Y-G, Chiappe LM, Meng Q-J, Li Q-G, Liu D. 2013. A new enantiornithine from the Yixian Formation with the first recognized avian enamel specialization. Journal of Vertebrate Paleontology 33(1):1-12. [featured article]

O’Connor JK, Zhou Z-H, Zheng X-T. Zheng et al. reply. 2013. Nature.

Wang Y-M, O’Connor JK, Li D-Q, You H-L. 2013. Previously unrecognized ornithuromorph bird diversity in the Early Cretaceous Changma Basin, Gansu Province, Northwestern China. PLoS ONE 8(10): e77693. doi:10.1371/journal.pone.0077693.

Wang M, O’Connor JK, Zhou Z-H. 2013. The first fossil crow (Corvus sp. indet.) from the Early Pleistocene Nihewan Paleolithic sites in North China. Journal of Archaeological Science 40(3): 1623-1628.

Zheng X-T, O’Connor JK, Huchzermeyer FH, Wang X-L, Wang Y, Wang M, Zhou Z-H. Exceptional preservation of ovarian follicles in Early Cretaceous birds and implications for early evolution of avian reproductive behaviour. 2013. Nature 495: 507-511. (corresponding author)

Zhou S, Zhou Z-H, O’Connor JK. 2013. Anatomy of Archaeorhynchus spathula from the Early Cretaceous. Journal of Vertebrate Paleontology 33(1): 141-152.

Zhou S, Zhou Z-H, O’Connor JK. 2013. A new piscivorous ornithuromorph from the Jehol Biota. Historical BiologyDOI: 10.1080/08912963.2013.819504.


O’Connor JK. 2012. A revised look at Liaoningornis longidigitrus (Aves). Vertebrata Palasiatica 50(1): 25-37.

O’Connor JK, Sun C-K, Xu X, Wang X-L, Zhou Z-H. 2012. A new species of Jeholornis

with complete caudal integument. Historical Biology 4(1): 29-41.

O’Connor JK, Chiappe LM, Chuong C-M, Bottjer DJ, You H-L. 2012. Homology and potential cellular and molecular mechanisms for the development of unique feather morphologies in early birds. Geosciences 2: 157-177.

Brocklehurst N, Upchurch P, Mannion PD, O’Connor JK. 2012. The Completeness of the Fossil Record of Mesozoic birds: Implications for Early Avian Evolution. PLoS ONE 7(6): e39056

Zheng Z-T, Wang X-L, O’Connor JK, Zhou Z-H. 2012. Insight into the early evolution of the avian sternum from juvenile enantiornithines (Aves: Ornithothoraces). Nature Communications 3:1116, DoI: 10.1038/ncomms2104. (corresponding author)

Zhou S, Zhou Z-H, O’Connor JK. 2012. A new basal beaked ornithurine bird from the Lower Cretaceous of western Liaoning, China. Vertebrata Palasiatica 50(1): 9-24.


O’Connor JK, Chiappe LM. 2011. A revision of enantiornithine (Aves: Ornithothoraces) skull morphology. Journal of Systematic Palaeontology 9(1): 135-147.

O’Connor JK, Zhou Z-H, Xu X. 2011. Additional specimen of Microraptor provides first evidence of dinosaurs preying on birds. Proceedings of the National Academy of Sciences 108(49):19662–19665.

O’Connor JK, Chiappe LM, Gao C-L, Zhao B. 2011. Anatomy of the Early Cretaceous bird Rapaxavis pani (Aves: Enantiornithes). Acta Palaeontologica Polonica 56(3): 463–475.

O’Connor JK, Zhou Z-H, Zhang F-C. 2011. A reappraisal of Boluochia zhengi (Aves: Enantiornithes) and a discussion of intraclade diversity in the Jehol avifauna, China. Journal of Systematic Palaeontology 9(1): 51-63.

Ji S-A, Atterholt J, O’Connor JK, Lamanna MC, Harris JD, Li D-Q, You H-L, Dodson P. 2011. A new, three-dimensionally preserved enantiornithine (Aves: Ornithothoraces) from Gansu Province, China. Zoological Journal of the Linnean Society 162: 201-219.


O’Connor JK, Chiappe LM, Gao K-Q. 2010. A new species of ornithuromorph (Aves: Ornithothoraces) bird from the Jehol Group indicative of higher-level diversity. Journal of Vertebrate Paleontology 30(2): 311-321 [featured article]

O’Connor JK, Dyke GD. 2010. Synonyms and the taxonomy of Mesozoic birds: the case of Sinornis and Cathayornis(Aves: Enantiornithes). Proceedings from the 7th meetingof the Society of Avian Palaeontology and Evolution.62(1): 7-20.

Wang X-R, O’Connor JK, Zhao B, Chiappe LM, Gao C-H, Cheng X-D. 2010. A new species of Enantiornithes (Aves: Ornithothoraces) based on a well-preserved specimen from the Yixian Formation of northwestern China. Acta Geologica Sinica 84(2): 247-256. (corresponding author)

You H-L, Atterholt J, O’Connor JK, Lamanna MC, Harris JD, Li D-Q. 2010. A second ornithuromorph from the Changma Basin, Gansu Province, northwestern China. Acta Palaeontologica Polonica 55(4): 617 – 625.


O’Connor JK, Wang X-R, Chiappe LM, Gao C-H, Meng Q-J, Cheng X-D, Liu J-Y. 2009. Phylogenetic support for a specialized clade of Cretaceous enantiornithine birds with information from a new species. Journal of Vertebrate Paleontology 29(1):188-204.

Close RA, Vickers-Rich P, Chiappe LM, O’Connor JK, Rich TH, Kool L. 2009. Earliest Gondwanan bird from the Cretaceous of southeastern Australia. Journal of Vertebrate Paleontology 29(2):616–619.

Tseng Z-J, O’Connor JK, Wang X-M, and Prothero D. 2009. The first Old World occurrence of the North American mustelid Sthenictis (Carnivora, Mammalia). Geodiversitas 31(4): 743 – 751.


O’Connor JK, Prothero D, Wang X-M, Li Q, Qiu Z-D. 2008. Magnetic stratigraphy of the Lower Pliocene Gaotege beds, Inner Mongolia. New Mexico Museum of Natural History and Science Bulletin 44:431-436.

Gao C-L, Chiappe LM, Meng Q-J, O’Connor JK, Wang X-R, Cheng X, Liu J. 2008. A new basal lineage of Early Cretaceous birds from China and its implications on the evolution of the avian tail. Palaeontology 51:775-791.


You H-L, Lamanna MC, Harris JD, Chiappe LM, O’Connor JK, Ji S-A, Lü J-C, Yuan C-X, Li D-Q, Zhang X, Lacovara KJ, Dodson P, Ji Q. 2006. A nearly modern amphibious bird from the Early Cretaceous of northwestern China. Science 312:1640-1643.

Zhang Z-H, Hou L-H, Hasegawa Y, O’Connor JK, Martin LD, Chiappe LM. 2006. The first Mesozoic heterodactyl bird from China. Acta Geologica Sinica (English Edition) 80:631-635.


You H-L, O’Connor JK, Chiappe LM, Ji Q. 2005. A new fossil bird from the Early Cretaceous of Gansu Province, northwestern China. Historical Biology 17:7-14.

Book Chapters

O’Connor JK, Chiappe LM, Bell A. 2011. Pre-modern birds: avian divergences in the Mesozoic. In: Dyke GJ, Kaiser G (eds), Living Dinosaurs: the evolutionary history of modern birds. UC Press. Pp. 39-116.

Sanz JL, Chiappe LM, O’Connor JK, Marugán-Lobón J, Chamero B. 2016. Aves. In: Poyato-Ariza FJ & Buscalioni AD (eds), Las Hoyas: A Cretaceous Wetland. A multidisciplinary synthesis after 25 years of research on an exceptional fossil Lagerstatte from Spain. Pfeil Verlag, Munich, Alemania. 262 pp. ISBN 978-3-89937-153-6

O’Connor JK. In press. The plumage of basal birds. In: Foth C, Rauhut O (eds), The evolution of feathers: a palaeontological perspective. Springer. 

Conference Abstracts

2003. O’Connor JK, Prothero DR, and Wang X-M. New specimen of the Miocene mustelid Sthenictis from the Miocene of Inner Mongolia, China. Journal of Vertebrate Paleontology 23(3):79A.

2003. O’Connor JK, Prothero DR, and Wang, X-M. New specimen of the Miocene mustelid Sthenictis from the Miocene of Inner Mongolia, China. Geological Society of America 
Abstracts with Programs.

2003. O’Connor JK, Prothero DR, and Wang, X-M. New specimen of the Miocene mustelid Sthenictis from the Miocene of Inner Mongolia, China. Southern California Conference on Undergraduate Research.

2003. Liter M, Prothero DR, and O’Connor JK. Land Mammals from the Middle Miocene Round Mountain Silt, Northeast of Bakersfield, California. Western Association of Vertebrate Paleontology.

2004. O’Connor JK, Prothero DR, Wang X-M, Li Q, Qiu Z-D. Magnetic stratigraphy of the Lower Pliocene Gaotege beds, Inner Mongolia. Western Association of Vertebrate Paleontology.

2004. O’Connor JK, Chiappe LM. New Enantiornithine Bird from the Early Cretaceous of Gansu Province, Northwestern China. Journal of Vertebrate Paleontology 24(3):99A.

2005. O’Connor JK, Chiappe LM, and Gao Q-K. A new fossil bird from the lower Cretaceous Jiufotang Formation, Liaoning Province, northwestern China. Journal of Vertebrate Paleontology 25(3): 97A.

2006. O’Connor JK, Chiappe LM, and Gao Q-K. A new fossil bird from the Lower Cretaceous Jiufotang Formation, Liaoning Province, northwestern China. 2ndInternational Palaeontology Congress: 442.

2006. O’Connor JK, Meng Q-J, Wang X-R, and Chiappe LM. Longirostrine enantiornithine birds: Information from a new Chinese specimen.  Journal of Vertebrate Paleontology 26(3): 106A.

2007. O’Connor JK, Meng Q-J, Chiappe LM, Wang X-R. Longirostrine enantiornithine birds; information from a new Chinese specimen. CalPaleo.

2008. O’Connor JK, Chiappe LM. A Review of the enantiornithine (Aves: Ornithothoraces) skull. 8th meeting of the Society of Avian Paleontology and Evolution.

2008. O’Connor JK, Chiappe LM, Gao C-L. A review of the Enantiornithine (Aves: Ornithothoraces) skull with information from a new species. CalPaleo.

2008. O’Connor JK, Chiappe LM. A review of the enantiornithine (Aves: Ornithothoraces) skull. Journal of Vertebrate Paleontology 28(3): 97A.

2009. O’Connor JK, Chiappe LM and Bell A. A comprehensive phylogeny of Mesozoic birds. Journal of Vertebrate Paleontology 29(3): 157A.

2010. O’Connor JK, Zhou ZH, Zhang FC. A reappraisal of Boluochia zhengi (Aves: Ornithothoraces). Chinese Society of Vertebrate Paleontology [in Chinese]

2010. O’Connor JK, Zhou ZH, Zhang FC. A new look at ‘old’ birds from the Jehol Fauna. Journal of Vertebrate Paleontology 30(3): 141A.

2011. O’Connor JK, Zhou ZH. The morphology and early evolution of the avian sternum. 9th Annual Meeting of the European Association of Vertebrate Palaeontologists, Heraklion, Crete, Greece.

2011. O’Connor JK, Zhou ZH, Xu X. Small theropod with bird in stomach indicates both lived in trees. Journal of Vertebrate Paleontology 31(3).

2011. O’Connor JK. Recent advances in Chinese Mesozoic bird taxonomy and systematics. 26thmeeting of the Chinese Society of Paleontology [in Chinese]

2011. O’Connor JK, Zhou ZH. The morphology and early evolution of the avian sternum. IV Congreso Latinoamerico Paleontologia de Vertebrados, San Juan, Argentina.  

2012. O’Connor JK. Dietary evolution in Mesozoic birds. 72nd annual meeting of the Society of Vertebrate Paleontology

2012. O’Connor JK. A brief overview of the current data on Enantiornithes (Aves: Ornithothoraces). 9th meeting of theSociety of Avian Paleontology and Evolution.

2013. O’Connor JK. Tail evolution in early birds: soft tissue and skeletal evidence from the Jehol. 10th Annual Meeting of the European Association of Vertebrate Palaeontologists, Villers-sur-mer, France.

2013. O’Connor JK. Tail evolution in early birds: soft tissue and skeletal evidence from the Jehol. 73rd annual meeting of the Society of Vertebrate Paleontology. 

2013. O’Connor JK. The histology of two Early Cretaecous female birds. 2ndInternational Symposium on Paleohistology.

2013. O’Connor JK. New information challenging the monophyly of Mesozoic Aves. International Conference on Evolutionary Ornithologyin memory of Evgeny Kurochkin, Moscow, Russia. 

2014. Zhou Z-H, O’Connor JK. Early evolution of birds: perspectives from new fossil discoveries in China. 26th International Ornithological Congress [symposium keynote, presenter: O’Connor]

2014. O’Connor JK. Evolution of the biologically modern bird: evidence from the Early Cretaceous Jehol Biota. 4th International Palaeontological Congress.[symposium organizer]

2014. O’Connor JK. Early evolution of birds: perspectives from new fossil discoveries in China. IV Simposio Paleontologia en Chile. [keynote speaker]

2014. O’Connor JK, Zhou Z-H. Earliest stages in the evolution of the modern avian skeleton: Archaeopteryxand the Jehol avifauna compared. 74th annual meeting of theSociety of Vertebrate Paleontology. [symposium]

2015. O’Connor JK, Atterholt J. Evolution of rapid growth in Ornithothoraces. 3rdInternational Symposium on Paleohistology.

2015. O’Connor JK, Choiniere JC. The early evolution of Aves. Biennial meeting of the Systematics Association. [symposium]

2015. O’Connor JK, Sullivan C, Zhou Z-H, Zheng X-T. Evolution and functional significance of derived sternal ossification patterns in ornithothoracine birds. 75thannual meeting of theSociety of Vertebrate Paleontology.

2016. O’Connor JK, Wang M, Hu H, Wang W, Zhou Z-H. A review of recent discoveries of ornithothoracines from the Cretaceous of China. 10th meeting of the Society of Avian Paleontology and Evolution.

2016. O’Connor JK. Paravian sternal ossification patterns. Meeting of the Chinese Society of Vertebrate Paleontology [keynote speaker]

2016. O’Connor JK, Zheng X-T, Wang X-L, Zhou Z-H. The biology of Eoconfuciusornis. 76th annual meeting of theSociety of Vertebrate Paleontology (book of abstracts p. 189).  

2017. O’Connor JK, Erickson G, Norell M, Hu H, Zhou Z-H. Definitive occurrence of medullary bone in an enantiornithine (Aves: Ornithothoraces). 4th International Symposium on Paleohistology.

2017. O’Connor JK. The trophic habits of early birds. 77th annual meeting of the Society of Vertebrate Paleontology.

2018. O’Connor JK, Zhou Z-H. Pygostyle formation in early birds. 5th International Palaeontological Congress.

2018. O’Connor JK, Maina J, Pan Y-H, Wang M, Wang X-L, Wang Y, Zheng X-T, Zhou Z-H. A specimen of Archaeorhynchuspreserving significant soft tissue including the first probable occurrence of fossilized lungs. 78th annual meeting of theSociety of Vertebrate Paleontology

2018. O’Connor JK. The trophic habits of early birds. 62nd annual meeting of thePalaeontological Society (invited speaker)

2019. O’Connor JK. Soft tissue preservation in fossil birds from the Early Cretaceous Jehol Biota. 4th International Meeting for Early-stage Researchers in Paleontology (Keynote speaker)

2019. O’Connor JK, Bailleul AM, Wang M, Zhou Z-H. Histology of juvenile enantiornithines from the Early Cretaceous Jehol Biota. 5th International Symposium on Paleohistology.

2019. O’Connor JK, Li D-P, Zheng X-T, Wang X-L, Wang Y, Zhou Z-H. Microraptor with ingested lizard preserved in situ and preliminary reconstruction of the Jehol food web. Geological Society of America Annual Meeting

2019. O’Connor JK, Bailleul AM, Wang M, Li Z-H, Zhou Z-H. Investigating probable reproductive tissues in stem birds using advanced microscopic analytical methods. 79th Annual Meeting of the Society of Vertebrate Paleontology (invited symposium speaker)

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